You find HoloMonitor systems in use all over the world. Some of them are listed below. Peer-reviewed articles and other publications discussing HoloMonitor applications and PHI’s core technology are available here.
Customer Publications
Reduced glutathione decreases cell adhesion and increases cell volume
Research Areas: Cell research
Utility of Novel Drug Targets for Treatment of Metastatic Cancer
Research Areas: Cancer Research
PIP5K1α is Required for Promoting Tumor Progression in Castration-Resistant Prostate Cancer
Research Areas: Cancer research
Self-Organization of Tissue Growth by Interfacial Mechanical Interactions in Multilayered Systems
Research Areas: Cell study
Chlorine exposure induces Caspase-3 independent cell death in human lung epithelial cells
Research Areas: Toxicology
Interfering with pak4 Protein Expression Affects Osteosarcoma Cell Proliferation and Migration
Research Areas: Cancer research
BRAFV600E induces reversible mitotic arrest in human melanocytes via microRNA-mediated suppression of AURKB
Research Areas: Cancer research
Digital holographic microscopy: a noninvasive method to analyze the formation of spheroids
Research Areas: Cancer research
Trametinib Inhibits the Growth and Aerobic Glycolysis of Glioma Cells by Targeting the PKM2/c-Myc Axis
Research Areas: Cancer Research
Mast cell tryptase enhances wound healing by promoting migration in human bronchial epithelial cells
Research Areas: Cell research
Breast cancer cell line toxicity of a flavonoid isolated from Baccharis densiflora
Research Areas: Cancer research
Effect of three-dimensional ECM stiffness on cancer cell migration through regulating cell volume homeostasis
Research Areas: Cancer research
IL4 promotes phagocytosis of murine leukemia cells counteracted by CD47 upregulation
Research Areas: Leukemia
Externally added cystatin C reduces growth of A375 melanoma cells by increasing cell cycle time
Research Areas: Cancer research
Evaluation of the Toxicity on Lung Cells of By-Products Present in Naphthalene Secondary Organic Aerosols
Research Areas: Aerosol toxicity
Epac1 signaling pathway mediates the damage and apoptosis of inner ear hair cells after noise exposure in a rat model
Research Areas: Hearing loss studies
Monobutyl phthalate can induce autophagy and metabolic disorders by activating the ire1a-xbp1 pathway in zebrafish liver
Research Areas: Chemical toxicity
Monoclonal antibody 2C5 specifically targets neutrophil extracellular traps
Research Areas: Cell Research
Targeted inhibition of ERα signaling and PIP5K1α/Akt pathways in castration‐resistant prostate cancer
Research Areas: Cancer research
Efficient and crucial quality control of HAP1 cell ploidy status
Research Areas: Cell Research
Smart drug combinations for cervical cancer: dual targeting of Bcl-2 family of proteins and aurora kinases
Research Areas: Cancer research
Exploiting the potential of commercial digital holographic microscopy by combining it with 3D matrix cell culture assays
Research Areas: Method Development
SUMOylation of PUM2 promotes the vasculogenic mimicry of glioma cells via regulating CEBPD
Research Areas: Cancer research
Improved Autophagic Flux in Escapers from Doxorubicin-Induced Senescence/Polyploidy of Breast Cancer Cells
Research Areas: Cancer research
Monobutyl phthalate (MBP) induces energy metabolism disturbances in the gills of adult zebrafish (Danio rerio)
Research Areas: Chemical toxicity
Subpopulations of Organoid-Forming Cells Have Different Motility
Research Areas: Organoid formation
The osteogenic commitment of CD271+CD56+ bone marrow stromal cells (BMSCs) in osteoarthritic femoral head bone
Research Areas: Stem cell therapy
Label-Free Classification of Apoptosis, Ferroptosis and Necroptosis Using Digital Holographic Cytometry
Research Areas: Basic Cell Research
Assembly of Epithelial Monolayers and Transmigration of Cancer Cells Captured with Phase Holographic Imaging
Research Areas: Cancer research
MicroRNA-362-3p Inhibits Migration and Invasion via Targeting BCAP31 in Cervical Cancer
Research Areas: Cancer research
Defining COMMD4 as an anti-cancer therapeutic target and prognostic factor in non-small cell lung cancer
Research Areas: Cancer research
Exosomes derived from mesenchymal stem cells repair a Parkinson’s disease model by inducing autophagy
Research Areas: Stem cell therapy
The human papillomavirus confers radiosensitivity in oropharyngeal cancer cells by enhancing DNA double strand break
Research Areas: Cancer research
Alkyl-carbon chain length of two distinct compounds and derivatives are key determinants of their anti-Acanthamoeba activities
Research Areas: Parasitic infections
N-terminal acetylation of actin by NAA80 is essential for structural integrity of the golgi apparatus
Research Areas: Cell Research
Antibacterial AZT derivative regulates metastasis of breast cancer cells
Research Areas: Cancer research
BCAP31, a cancer/testis antigen-like protein, can act as a probe for non-small-cell lung cancer metastasis
Research Areas: Cancer research
Berberine Maintains the Neutrophil N1 Phenotype to Reverse Cancer Cell Resistance to Doxorubicin
Research Areas: Frontiers in Pharmacology
LRIG1 in lung cancer: prognostic effects and mechanistic studies
Research Areas: Cancer Research
Development of label-free biophysical markers in osteogenic maturation
Research Areas: Cell Research
KLF2+ stemness maintains human mesenchymal stem cells in bone regeneration
Research Areas: Stem cell therapy
Synthesis of a dihalogenated pyridinyl silicon rhodamine for mitochondrial imaging by a halogen dance rearrangement
Research Areas: Flourofore synthesis
Propofol prevents oxidative stress and apoptosis by regulating iron homeostasis and targeting JAK/STAT3 signaling in SH-SY5Y cells
Research Areas: Neurodegenerative disorders
Unravelling the role of trophoblastic-derived extracellular vesicles in in regulatory T cell differentiation
Research Areas: Cell Research
Exploring the Natural Piericidins as Anti-Renal Cell Carcinoma Agents Targeting Peroxiredoxin 1
Research Areas: Cytotoxicity
The long noncoding RNA TP73‐AS1 promotes tumorigenicity of medulloblastoma cells
Research Areas: Cancer research
ITGA2 as a potential nanotherapeutic target for glioblastoma
Research Areas: Cancer research
In vitro SOD-like activity of mono- and di-copper complexes with a phosphonate substituted SALAN-type ligand
Research Areas: Cytotoxicity
Effects of GADL1 overexpression on cell migration and the associated morphological changes
Research Areas: Bipolar disorder treatment
Whole-Cell Multiparameter Assay for Ricin and Abrin Activity-Based Digital Holographic Microscopy
Research Areas: Cytotoxicity
Digital Holographic Cytometry: Macrophage Uptake of Nanoprobes
Research Areas: Cytotoxicity
Emodin regulates neutrophil phenotypes to prevent hypercoagulation and lung carcinogenesis
Research Areas: Cancer research
Preparation of fibrin hydrogels to promote the recruitment of anti-inflammatory macrophages
Research Areas: Tissue regeneration
Metabolism of diethyl phthalate (DEP) and identification of degradation intermediates by Pseudomonas sp. DNE-S1
Research Areas: Biodegradation
Hippo pathway controls cell adhesion and context-dependent cell competition to influence skin engrafment efficiency
Research Areas: Skin engraftment
The photocytotoxicity effect of cationic sulfonated corrole towards lung cancer cells: in vitro and in vivo study
Research Areas: Cancer research
Moving into a New Dimension: Tracking Migrating Cells with Digital Holographic Cytometry in 3D
Research Areas: Cancer research
Nuclear respiratory factor 1 acting as an oncoprotein drives estrogen-induced breast carcinogenesis
Research Areas: Cancer research
Lack of LTβR Increases Susceptibility of IPEC-J2 Cells to Porcine Epidemic Diarrhea Virus
Research Areas: Cell research
The connection between the cardiac glycoside‐induced senescent cell morphology and Rho/Rho kinase pathway
Research Areas: Cancer research
Evaluation of Holographic Imaging Cytometer HoloMonitor M4 Motility Applications
Research Areas: Cancer research
Single cell analysis of proliferation and movement of cancer and normal-like cells on nanowire array substrates
Research Areas: Nanomaterials and cell interactions
Ulleungdin, a lasso peptide with cancer cell migration inhibitory activity discovered by the genome mining approach
Research Areas: Cancer Research
TRPM2 Modulates Neutrophil Attraction to Murine Tumor Cells by Regulating CXCL2 Expression
Research Areas: Immunoncology
Hexagonal Boron Nitride Functionalized with Au Nanoparticles—Properties and Potential Biological Applications
Research Areas: Biocompatibility
General Cytotoxicity and Its Application in Nanomaterial Analysis
Research Areas: Nanomateiral
Bi-allelic Loss of CDKN2A Initiates Melanoma Invasion via BRN2 Activation
Research Areas: Melanoma
Quantitative Phase Imaging for Label-Free Analysis of Cancer Cells—Focus on Digital Holographic Microscopy
Research Areas: Digital holography technology
Quantitative evaluation of morphological changes in activated platelets in vitro using digital holographic microscopy
Research Areas: Platelet activation
The impact of circulating preeclampsia-associated extracellular vesicles on the migratory activity and phenotype of THP-1 monocytic cells
Research Areas: Pathogenesis in preeclampsia
The evolution of phase holographic imaging from a research idea to publicly traded company
Research Areas: Quantitative phase microscopy
Quantification of mammalian tumor cell state plasticity with digital holographic cytometry
Research Areas: Melanoma
Applications of quantitative time lapse holographic imaging to the development of complex pharmaceutical nano formulations
Research Areas: Method development
A Time-lapse, Label-free, Quantitative Phase Imaging Study of Dormant and Active Human Cancer Cells
Research Areas: Cancer Research
β-Nitrostyrene 1 derivatives attenuate LPS-mediated acute lung injury via the 2 inhibition of neutrophil-platelet interactions and NET release
Research Areas: Acute lung injury (ALI) and the acute respiratory distress syndrome (ARDS)
Tau Phosphorylation and Amyloid-β Deposition in the Presence of Formaldehyde
Research Areas: Alzheimer’s disease
NatD promotes lung cancer progression by preventing histone H4 serine phosphorylation to activate Slug expression
Research Areas: Cancer research
The role of Vimentin in Regulating Cell Invasive Migration in Dense Cultures of Breast Carcinoma Cells
Research Areas: Cancer research
A Biophysical Approach to Quantify Skeletal Stem Cells Trans-differentiation as a Model for the Study of Osteoporosis
Research Areas: Stem cell research
Interleukin-6 blockade attenuates lung cancer tissue construction integrated by cancer stem cells
Research Areas: Cancer research
CFTR supports cell death through ROS-dependent activation of TMEM16F (anoctamin 6)
Research Areas: Cystic fibrosis
Synthesis, Characterization and in Vitro and in Vivo Photodynamic Activities of a Gallium(III) Tris(Ethoxycarbonyl)corrole
Research Areas: Photo-cytotoxicity; photodynamic therapy (PDT)
Hypoxia-inducible Factor-1 Alpha Is Involved in Rip-induced Necroptosis Caused by in Vitro and in Vivo Ischemic Brain Injury
Research Areas: Ischemic brain injury
Hydroxylated Fluorescent Dyes for Live-cell Labeling: Synthesis, Spectra and Super-resolution Sted
Research Areas: Method development
Label-free High Temporal Resolution Assessment of Cell Proliferation Using Digital Holographic Microscopy
Research Areas: Digital holography technology
Characterization of a Transgenic Mouse Model Exhibiting Spontaneous Lung Adenocarcinomas With a Metastatic Phenotype
Research Areas: Cancer research
The “inherent Vice” in the Anti-angiogenic Theory May Cause the Highly Metastatic Cancer to Spread More Aggressively
Research Areas: Cancer research
Promising Anticancer Activity of Cyclotrichium Niveum L. Extracts Through Induction of Both Apoptosis and Necrosis
Research Areas: Cancer research
Characterization of Dormant and Active Human Cancer Cells by Quantitative Phase Imaging
Research Areas: Angiogenesis
Holography: the Usefulness of Digital Holographic Microscopy for Clinical Diagnostics
Research Areas: Holography technology
Loss of Erα Induces Amoeboid-like Migration of Breast Cancer Cells by Downregulating Vinculin
Research Areas: Cancer research
Three-dimensional Morphology of C6/36 Cells Infected by Dengue Virus: a Study Based on Digital Holographic Microscopy
Research Areas: Virus infection
Supervised Classification of Etoposide-treated in Vitro Adherent Cells Based on Noninvasive Imaging Morphology
Research Areas: Image-based analysis, cell classification
Cathelicidins Positively Regulate Pancreatic Β-cell Functions
Research Areas: Immunregulation
Ca2+ signals, cell membrane disintegration, and activation of tmem16f during necroptosis
Research Areas: Cell death
Relationship Between tmem16a/anoctamin 1 and lrrc8a
Research Areas: Basic cell research
Shielding of the Geomagnetic Field Alters Actin Assembly and Inhibits Cell Motility in Human Neuroblastoma Cells
Research Areas: Mode of action
Enhanced Cytotoxicity of Folic Acid-targeted Liposomes Co-loaded With C6 Ceramide and Doxorubicin: in Vitro Evaluation on HeLa, A2780-ADR and H69-AR Cells
Research Areas: Cancer research, Drug development
See in 3D: state of the art of 3D display technologies
Research Areas: Digital holography technology
Macrophage Polarization and the Effect of Microrna-155 Administered in Water-in-oil-in-water Multiple Emulsion Formulations
Research Areas: Cancer research, immunology
Interfacing Antibody-based Microarrays and Digital Holography Enables Label-free Detection for Loss of Cell Volume
Research Areas: Cancer research, drug development
Induction of Morphological Changes in Death-induced Cancer Cells Monitored by Holographic Microscopy
Research Areas: Cell research
Environmental and Toxicological Impacts of Glyphosate With Its Formulating Adjuvant
Research Areas: Drug development
Nanowires in Cell Biology – Exploring Interactions and Applications
Research Areas: Nanowires
Digital Holographic Microscopy for Non-invasive Monitoring of Cell Cycle Arrest in L929 Cells
Research Areas: Method development
Edaravone Protects Against Methylglyoxal-induced Barrier Damage in Human Brain Endothelial Cells
Research Areas: Drug development
Protein kinase A (PKA) pathway is functionally linked to androgen receptor (AR) in the progression of prostate cancer
Research Areas: Cancer research
Fibroblasts Cultured on Nanowires Exhibit Low Motility, Impaired Cell Division, and Dna Damage
Research Areas: Nanomaterials
Cells and Holograms — Holograms and Digital Holographic Microscopy as a Tool to Study the Morphology of Living Cells
Research Areas: Digital holographic technology
A Unifying Mechanism for Cancer Cell Death Through Ion Channel Activation by Hamlet
Research Areas: Action of HAMLET
Prevention and Treatment of Colon Cancer by Peroral Administration of Hamlet
Research Areas: Action of HAMLET
Low Resolution Solution Structure of Hamlet and the Importance of Its Alpha-domains in Tumoricidal Activity
Research Areas: Action of HAMLET
HAMLET: Functional Properties and Therapeutic Potential
Research Areas: Action of HAMLET
Dermatan Sulfate Is Involved in the Tumorigenic Properties of Esophagus Squamos Cell Carcinom
Research Areas: Cancer research
HAMLET Binding to Α-actinin Facilitates Tumor Cell Detachment
Research Areas: Action of HAMLET
Generation of Trisomies in Cancer Cells by Multipolar Mitosis and Incomplete Cytokinesis
Research Areas: Cancer research
Digital Holographic Microscopy — Innovative and Non-destructive Analysis of Living Cells
Research Areas: Digital holographic technology
Cell motility studies using digital holographic microscopy
Research Areas: Digital holographic technology
Non-invasive, Label-free Cell Counting and Quantitative Analysis of Adherent Cells Using Digital Holography
Research Areas: Digital holographic technology
Design of Reconfigurable Hardware Architectures for Real-Time Applications
Research Areas: Digital holography technology
A Hardware Acceleration Platform for Digital Holographic Imaging
Research Areas: Digital holographic technology
Modeling and Exploration of a Reconfigurable Architecture for Digital Holographic Imaging
Research Areas: Background technology
Object Characterization With Refractometric Digital Fourier Holography
Research Areas: Digital holographic technology
High-resolution Digital Transmission Microscopy — a Fourier Holography Approach
Research Areas: Digital holographic technology
Refractometry of Microscopic Objects With Digital Holography
Research Areas: Digital holography technology
AUSTRALIA
- Monash University, Melbourne
- Bosch Institute, University of Sydney, Sydney
- The Science Place, James Cook University, Townsville
- QIMR Berghofer Medical Research Institute, Brisbane
- Menzies Health Institute, Griffith University, Gold Coast
- Vascular Biology Unit, James Cook University, Townsville
- Institute of Health and Biomedical Innovation, Queensland University of Technology, Brisbane
- Queensland Centre for Drug Target Screening and Characterisation, Queensland University of Technology, Brisbane
Belgium
- IRIBHM, Université Libre de Bruxelles
Canada
- Bluerock Therapeutics, Toronto
- Cardiovascular Research Centre, University of Alberta, Edmonton
CHINA
- Institute of Pharmacy, Henan University, Kaifeng
- Biosafety Level 3 Laboratory, Southern Medical University, Guangzhou
- Department of Immunology, The Fourth Military Medical University, Xi’an
- State Key Laboratory of Proteomics, Institute of Radiation Medicine, Beijing
- Department of Pharmaceutical Sciences, Binzhou Medical University, Yantai
- Institute of Animal Science, Chinese Academy of Agricultural Sciences, Beijing
- Institute of Biomechanics and Medical Engineering, Tsinghua University, Beijing
- School of Resources and Environment, Northeast Agricultural University, Harbin
- Laboratory of Molecular Iron Metabolism, Hebei Normal University, Shijiazhuang
- State Key Laboratory of Food Science and Technology, Jiangnan University, Wuxi
- Department of Neurology, First Hospital of Hebei Medical University, Shijiazhuang
- The State Key Laboratory of Pharmaceutical Biotechnology, Nanjing University, Nanjing
- Department of Neurosurgery, Shengjing Hospital of China Medical University, Shenyang
- The Affiliated Stomatologic Hospital, School of Medicine, Zhejiang University, Hangzhou
- State Key Laboratory of Brain and Cognitive Science, Chinese Academy of Sciences, Beijing
- Department of Biochemistry and Molecular Biology, Guangdong Medical University, Zhanjiang
- Tianjin Key Laboratory of Neurotrauma Repair, University of Chinese People’s Armed Police Force, Tianjin
- Molecular Medicine Research Center, Sichuan University and Collaborative Innovation Center, Chengdu
- School of Biomedical and Pharmaceutical Sciences, Guangdong University of Technology, Guangzhou
- Institute of Sports Medicine, Shandong First Medical University & Shandong Academy Medical Sciences, Shandong
- State Key Laboratory of Bioactive Substances and Functions of Natural Medicines, Chinese Academy of Medical Sciences and Peking Union Medical College, Beijing
CZECH REPUBLIC
- Faculty of Medicine and Dentistry, Palacký University Olomouc
Best microscope I’ve tried so far — worth every penny!
Markus Bosteen
Senior Scientist, Novo Nordisk, Copenhagen
Denmark
- Novo Nordisk A/S
FRANCE
- Genethon
- Claude Bernard University, Lyon
- IMS Laboratory, University of Bordeaux
- Department of Biology, University of Le Havre
GERMANY
- Bayer A/G
- Evotec International GmbH, Göttingen
- Institute of Phycology, University of Regensburg
- Centre for Biological Signalling Studies, University Hospital Freiburg
- GSI Helmholtzzentrum für Schwerionenforschung GmbH, Darmstadt
- Light Microscopy Facility, German Cancer Research Center (DKFZ)
- Department Physiology and Pathology of Ion Transport, Leibniz-Institut für Molekulare Pharmakologie
HUNGARY
- Avidin Biotechnology
- The Research Institute of Technical Physics and Materials Science
- Department of Genetics, Cell- and Immunobiology, Semmelweis University
Our [HoloMonitor] microscope lives directly in our cell culture incubator and allows us to see how various treatments affect our cells in real-time with three-dimensional single-cell resolution — it’s amazing!
Alexander Payumo
San Jose State University
ISRAEL
- Israel Institute for Biological Research (IIBR)
- Diabetes Research Center, The Hebrew University of Jerusalem
- Department of Life Sciences, Ben-Gurion University of the Negev, Beer Sheva
- Raymond and Beverly Sackler School of Physics and Astronomy, Tel Aviv University, Tel Aviv
ITALY
- Institute of Biophysics, National Research Council of Italy, Genoa
- Department of Civil Engineering and Architecture, University of Pavia
JAPAN
- Laboratory of Biomaterials, Kyoto University, Kyoto
- Division of Oral Bioengineering, Niigata University, Niigata
- Department of Molecular Medicine and Biochemistry, Akita University
- Faculty of Pharmacy Advanced Biomedical Products, Kindai University
- Department of Biochemistry and Molecular Biology, Kobe University, Kobe
- Center for Stem Cell Biology and Regenerative Medicine, University of Tokyo
KOREA
- Division of Life Sciences, Korea University, Seoul
- Chemical Biology Research Center, Korea Research Inst. of Bioscience and Biotechnology (KRIBB)
As a label-free, non-toxic imaging system, HoloMonitor M4 provides us the means to begin investigating the immediate reaction of melanoma cells to acute environmental and genetic stimuli.
DR ROBERT JUDSON-TORRES
UNIVERSITY OF UTAH
Lithuania
- Lithuanian Center for Innovative Medicine
MALAYSIA
- Institute for Research in Molecular Medicine, Universiti Sains Malaysia
Netherlands
- Biophysical chemistry, Radboud University
NEW ZEALAND
- University of Auckland, Auckland

NORWAY
- Institute of Clinical Medicine, University of Oslo
- Department of Biomedicine, University of Bergen
POLAND
- Institute of Medical Biology, Polish Academy of Sciences, Lodz
- Department of Physiology, Pomeranian Medical University in Szczecin
- Department of Microbiology and Immunology, Medical University of Silesia
- Department of Biology and Pharmaceutical Botany, Medical University of Gdańsk
- Department of Clinical Immunology, Jagiellonian University Medical College, Krakow
RUSSIA
- InnovaBio
SINGAPORE
- Esco Micro Pte. Ltd.
We are very happy with the HoloMonitor M4. It is very easy to use, produces consistent results since nearly 4 years and is a great way to gather a lot of quantitative biophysical metrics on adherent cells in a label-free and real-time way.
PIERRE BAGNANINCHI
THE UNIVERSITY OF EDINBURGH
SWEDEN
- Fluicell AB
- Cline Scientific AB
- Solid State Physics, Lund University
- Health and Society, Malmö University
- Department of Biology, Lund University
- Sahlgrenska University Hospital, Gothenburg
- Department of Molecular Biology, Umeå University
- Clinical Chemistry and Pharmacology, Lund University
- Department of Experimental Medical Science, Lund University
- Nursing and Medical Technology, Luleå University of Technology
- Sahlgrenska Center for Cancer Research, University of Gothenburg
Switzerland
- University of Fribourg
TAIWAN
- Institute of Biomedical Sciences, Academia Sinica, Taipei
- Laboratory Animal Center, Taipei Medical University, Taipei
- Graduate Institute of Biomedical Sciences, Chang Gung University, Taoyuan
- Graduate Institution of Biomedical and Pharmaceutical Science, Fu Jen Catholic University, New Taipei City
TURKEY
- Department of Biochemistry, Ege University
- Department of Biochemistry, Dumlupınar University
The HoloMonitor M4 is a beneficial device for live-cell imaging, providing a lot of data. The device does not require large space and the software application is user-friendly, while recorded pictures look great. The good thing is that you can analyze your experiment using one software assay, and later, you can rerun the recorded data in another assay, which allows you to get the maximum from your experiment.
Sasa Vasilijic, PhD
UK
- University of Aberdeen
- Imperial College, London
- St James´s University Hospital, Leeds
- Bristol Medical School, University of Bristol
- Institute of Aquaculture, University of Stirling
- School of Engineering, University of Glasgow
- Department of Chemical Engineering, Loughborough University
- MRC Centre for Regenerative Medicine, The University of Edinburgh
- Institute of Infection, Immunity & Inflammation, University of Glasgow
- School of Health and Life Sciences, University of West Scotland, Glasgow
- Healthcare Science Research Institute, Manchester Metropolitan University
US
- Artemys Foods
- Gladstone Institutes
- Pandorum Technology
- Johns Hopkins University
- Penn State College of Medicine
- Biological Sciences, San Jose State University
- Huntsman Cancer Institute, University of Utah
- OHNS/Otology & Neurotology, Stanford University
- Pharmaceutical Sciences, Northeastern University
- School of Pharmacy, North Dakota State University
- Vascular Biology Program, Boston Children’s Hospital
- Bioengineering, University of Massachusetts Dartmouth
- National Institute on Aging, National Institutes of Health
- Harvard Medical School, Partners HealthCare International
- Wake Forest Institute of Regenerative Medicine, Winston-Salem
- Cell Biology and Human Anatomy, University of California, Davis
- Department of Ophthalmology and Visual Sciences, University of Iowa
- Cardiovascular Research Institute, University of California, San Francisco